At the Netherfield Ball, Elizabeth Bennet and Mr. Darcy are at wit’s end with each other. Darcy can no longer deny his growing attraction to her, but she is clearly not ready to hear him. Elizabeth is confused by the many versions of Mr. Darcy she has encountered. She seeks refuge from the crowds, including her ridiculous family, on the terrace. He finds solace in the empty library. Grappling with their own feelings, each look up at the night sky. The evening is fine and the stars shine in their multitude reminding Elizabeth and Darcy just how small they are. While looking up at the Heavens, each notices a shooting star streaking across the sky. And each makes a wish: to better understand the other. Neither is ready for what happens next.

The day dawns bright but almost immediately, Elizabeth realizes something is very wrong. She is not in her room, not even in her own home. But it was not until she saw her - or rather his - reflection in the mirror that the magnitude of her situation becomes apparent. She has somehow come to inhabit Mr. Darcy’s body! Meanwhile Mr. Darcy, who had imbibed too much wine the night before, remains blissfully unaware of his situation while he sleeps. It was not until much later in the day than is his usual wont, that he finally awakes and discovers that he too is not himself.

While Through Each Other's Eyes stays true to the overall character of the Pride and Prejudice, the radical change in the characters’ situation allows for some changes in perspective and timing, as well as some sensual situations but no explicit sex scenes. It includes all the original characters but also adds a few new ones.