In the wake of a bloody battle at Forward Operating Base Frontier and a scandalous presidential impeachment, Lieutenant Colonel Jan Thorsson, call sign " Harlequin," becomes a national hero and a pariah to the military that is the only family he' s ever known. In the fight for Latent equality, Oscar Britton is positioned to lead a rebellion in exile, but a powerful rival beats him to the punch: Scylla, a walking weapon who will stop at nothing to end the human-sanctioned apartheid against her kind.

When Scylla' s inhuman forces invade New York City, the Supernatural Operations Corps are the only soldiers equipped to prevent a massacre. In order to redeem himself with the military, Harlequin will be forced to face off with this havoc-wreaking woman from his past, warped by her power into something evil.