Of all the Legiones Astartes, the White Scars of Jaghatai Khan remain the most enigmatic and elusive. Born of a civilisation that prizes honour, speed and fearsome loyalty, their allegiance has yet remained unclear even as the galaxy is torn apart by Horus's treachery, and both sides have apparently counted them among their potential allies in the war to come.

In the wake of the Dropsite Massacre at Isstvan V, the survivors of the Salamanders Legion searched long and hard for their fallen primarch, but to no avail. Little did they know that while Vulkan might have wished himself dead, he lives still...languishing in a hidden cell for the entertainment of a cruel gaoler, his brother Konrad Curze.

Once the brightest star in the Imperium and always first among his primarch brothers, Horus has dragged the Space Marine Legions into the bloodiest conflict that the galaxy has ever seen. While their allies wage war on a thousand different fronts, the XVIth Legion descend upon the Knight world of Molech - home to the ruling House Devine, and a principle stronghold of the Imperial Army.

The Heresy came to Calth without warning. In just a few hours of betrayal and bloodshed, the proud warriors of the XIIIth Legion - Guilliman's own Ultramarines - were laid low by the treachery of their erstwhile brothers of the XVIIth. Now, as the planet is scoured by solar flares from the wounded Veridian star, the survivors must take the fight to the remaining Word Bearers and their foul allies, or face damnation in the gloomy arcology shelters beneath the planet's surface.

The Shadow Crusade has begun. While the Ultramarines reel from Kor Phaeron's surprise attack on Calth, Lorgar and the rest of the Word Bearers strike deep into the realm of Ultramar. Their unlikely allies, Angron and the World Eaters, continue to ravage each new system they come across - upon the garrison planet of Armatura, this relentless savagery may finally prove to be their undoing. Worlds will burn, Legions will clash and a primarch will fall.

In the aftermath of the Dropsite Massacre at Isstvan V, a battered and bloodied force of Iron Hands, Raven Guard and Salamanders regroups on a seemingly insignificant death world. Fending off attacks from all manner of monstrous creatures, the fractious allies find hope in the form of human refugees fleeing from the growing war and cast adrift upon the tides of the warp.

Vulkan lies in state beneath the Fortress of Hera, and yet many of his sons still refuse to believe that he is truly dead. After a seemingly miraculous rescue by the Ultramarines, Artellus Numeon, once captain of the Pyre Guard, urges the other Salamanders on Macragge to leave Imperium Secundus and return their primarch's body to the home world of Nocturne - there to be reborn in the flames of Mount Deathfire.

Perturabo - master of siegecraft and the executioner of Olympia. Long has he lived in the shadow of his more favoured primarch brothers, frustrated by the mundane and ignominious duties which regularly fall to his Legion. When Fulgrim offers him the chance to lead an expedition in search of ancient and destructive xenos weaponry, the Iron Warriors and the Emperor's Children unite and venture deep into the heart of the great star maelstrom that haunts Perturabo's dreams.

Since the earliest days of the Great Crusade, Sanguinius - angelic primarch of the IXth Legion - was ever among the closest and most loyal of Horus' brothers. But the Blood Angels have long kept their true nature hidden from the rest of the Imperium, and when the Warmaster hints that the key to their salvation may lie in the ruins of a conquered world, the sons of Sanguinius race to claim it.

With the Dark Angels spread across a hundred systems, primarch Lion El'Jonson stands as Lord Protector of Ultramar - though his true motives are known to few indeed, and old rivalries on the home world threaten to tear the Legion in half. But when word comes of the Night Lords' attack on Sotha, the Lion's brutal actions bring Imperium Secundus once again to the brink of civil war.

Unaware of the wider Heresy and following the Warmaster's increasingly cryptic orders, Roboute Guilliman returns to Ultramar to muster his Legion for war against the orks massing in the Veridian system. Without warning, their supposed allies in the Word Bearers Legion launch a devastating invasion of Calth, scattering the Ultramarines fleet and slaughtering all who stand in their way. This confirms the worst scenario Guilliman can imagine - Lorgar means to settle their bitter rivalry once and for all.

Recalled from the Great Crusade after Ullanor, Rogal Dorn and the VIIth Legion were appointed as the Emperor's praetorians - but only after the Warmaster's treachery was revealed did the full extent of that sacred duty become apparent. Now, the Solar System comes under attack for the first time since the war began, and many of the seemingly impregnable defences wrought by the Imperial Fists prove inadequate. With all eyes fixed firmly upon this new threat beyond the gates of Terra, who in turn will protect Dorn from the enemy within?

The Emperor's vision of mankind ascendant lies in tatters. The Great Crusade is naught but a fading memory. The dream of Unity is no more. But with Horus's rebellion spreading to every corner of the Imperium and war engulfing new worlds and systems almost daily, there are some who now ask: were the signs there to be seen all along?

For too long had the Vth Legion ranged out beyond the sight of the wider Imperium, remaining ignorant of the Warmaster's rebellion and the war that inevitably followed. Only once their primarch, Jaghatai Khan, had satisfied himself that the path before them was just and true did the White Scars choose a side, taking the fight to the traitors on every front.

The galaxy burns at Horus' command, and a billion more lives are lost with each passing moment - though it was not always so. Once, the title of Warmaster stood for honour, loyalty and a fierce pride in the strength of the Space Marine Legions. But perhaps by following the myriad lines of destiny and defiance that were already woven around the primarchs and their sons, we may yet come to understand the bitterness that can eat away at even the most steadfast of souls....