Mma Ramotswe and Mma Makutsi are called to a safari lodge in Botswana’s Okavango Delta to carry out a delicate mission on behalf of a former guest. The Okavango makes Precious appreciate once again the beauty of her homeland: it is a paradise of teeming wildlife, majestic grasslands and sparkling water. However, it is also home to rival safari operators, fearsome crocodiles and disgruntled hippopotamuses.

As the countdown to Mma Makutsi’s wedding begins, all is not as it should be at the No.1 Ladies’ Detective Agency. While investigating unpleasant occurrences on a southern cattle-post, Mma Ramotswe, always on the side of the weak against the strong, has reason to reflect on Rule No.3 of The Principles of Private Detection: never lie to the client. Apprentice mechanic Charlie seems to be avoiding certain important responsibilities.

Precious Ramotswe, traditionally-built proprietor of Botswana's No. 1 Ladies' Detective Agency, contemplates the two sorts of problems that we all face in life: the major ones (such as drought), about which very little can be done, and the other sort which people make for themselves through human weakness and an inability to resist temptation. Like the inability to resist cake, for instance.

Precious Ramotswe, that cheerful Botswanan private investigator of 'traditional build', is now married to Mr J.L.B. Matekoni of Tlokweng Road Speedy Motors. The Agency is busy, but Mma Ramotswe cannot ignore the plea made by a woman who comes to her with a tale of particular misfortune. Unfortunately, her attempts to help are interrupted by a close encounter between her tiny white van and a bicycle.

Mr J.L.B. Matekoni is also drawn into the work of the agency when he investigates a case involving an errant husband. But can a man investigate such matters with as much skill as one of the ladies? Mma Ramotswe has her doubts. One thing, though, that she does not doubt is the good nature of Mr J.L.B. Matekoni, who stands for all that is solid and true in a shifting world.

Mma Ramotswe, who became engaged to Mr J.L.B. Matekoni at the end of the first audiobook, is still engaged. She wonders when a day for the wedding will be named, but she is anxious to avoid putting too much pressure on her fiancé. For indeed he has other things on his mind, notably a frightening request made of him by Mma Potokwani, pushy matron of the Orphan Farm.

Unusual requests are commonplace at the No. 1 Ladies' Detective Agency, but so are very ordinary ones, such as a plea for money: when a distant cousin calls on Mma Ramotswe to ask for help with another relative's hip operation, tricky questions arise in regard to family responsibility. Mma Makutsi and Mr J. L. B Matekoni are far from convinced of the merits of the request, but it is hard for Mma Ramotswe, with a heart her size, to turn people away.

Mma Ramotswe and Mma Makutsi have always relied on the advice of the classic guide to their trade, The Principles of Private Detection. But who is the eminent author, Mr Clovis Andersen, and what if he was to come to Botswana? That seems a very unlikely possibility, and yet… When Mr Andersen visits Botswana on holiday, he is delighted and intrigued to stumble across a roadside sign that reveals the existence of the No.1 Ladies' Detective Agency.

As Botswana awaits the familiar blessing of the rains and their eternal cycle, upheaval is taking place at the No.1 Ladies' Detective Agency. Not only is Mr J. L. B. Matekoni attempting to reform himself into a modern husband but, following her marriage to Phuti Radiphuti, Mma Makutsi has joyful news.With the arrival of an heir to the Double Comfort furniture empire and Mma Makutsi busy with motherhood, cobras and a senior aunt, Mma Ramotswe must grapple alone with tea-making, detective work and a new uncertainty.

Cephas Buthelezi certainly talks the talk, Precious discovers, but would he have the wherewithal to deal with her current case, a man who has been attacked by ostrich rustlers and is eager to reassess his life? Meanwhile, there are difficulties at the Tlokweng Road Speedy Motors, where one of the apprentices has discovered the Lord; problems at home with the mysterious death of a hoopoe; and romantic complications when Mma Makutsi sets up a typing school for men.