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Still haunted by early memories of growing up in an abusive family, 17-year-old Moon can't separate himself from his past. Twelve-year-old Harper comes from a world full of love and support. Despite their differences, they become the best of friends. Just prior to World War II, they spend the summer in the wilds of Washington State, with reclusive storekeeper and mountaineer, Olinger. But when winter comes, a blustery snowstorm hits, and Moon is missing.

The hero of this book is a prune. His journey begins in an ordinary way: he is picked, packed in a box with the rest of the crop, and carried off to the packing shed. But along the way, his life takes an unexpected turn. When his crate tips over, Prune meets Pica, a magpie. As they travel together, they meet Muskrat. The three soon become friends and begin an epic quest that will teach them much about loyalty, adventure, and courage.